Source areas of the Eastern Ebro Valley loess (NE Iberian Peninsula): Heavy mineral composition as a provenance indicator

Loess deposits of the Eastern Ebro Valley (NE Iberian Peninsula) occupy an area of ~2000 km2 mostly on the east of the Ebro depression and locally in the Móra Basin. We studied 38 samples of loess and 47 samples of their possible source areas, in order to determine the location of the origin of the loess materials, based on their particle size distribution and composition of heavy minerals. Our results clearly differentiate two depositional loess basins disconnected by the reliefs of the Pre‐Coastal Range with different mineralogical signatures: the Ebro Basin and the Móra Basin. The most relevant variables to discern source areas were the percentage of heavy minerals, the concentrations of zircon, staurolite, rutile and inosilicates, and the presence of gypsum. In the Ebro Basin, the largest set of loess deposits is located around a depocenter in the Batea region and along several lateral areas, with materials that come from the central alluvial plain of the Middle Ebro River (25 to >100 km travel distance), in addition to deflated materials from Miocene interfluvial substrates. On the other hand, loess deposits of the Móra Basin come from more proximal source areas (<15 km), originating from the alluvial materials of the Lower Ebro River that contain the contributions of its tributaries, the Cinca and Segre rivers. Taking into account the homogeneity in the mineralogical composition of complex loess sequences, we deduced a general WNW–ESE wind direction, which would have remained constant over the last two cold periods (MIS‐2 and MIS‐6). Along this axis, the Pàndols–Cavalls Range, with a height difference of 200–350 m, acted as a wind barrier separating the loess of Ebro Basin from the loess of Móra Basin.

rivers (Rhine, Danube, Rhône, Po), have their sources in the Alpine glacial system. In addition to the ice sheets and Alpine system, fluvially transported material coming from low mountain ranges also plays a major role in loess formation. Besides the glacier and fluvial-fed systems, other less well-developed loess areas appear at the periphery of aeolian sandsheets and dunefields, particularly along the Atlantic coasts (coastal systems) and also inland, in association with easily erodible detrital rocks such as sandstones, where strong winds are generated by relief (continental systems) (Bertran et al., 2016(Bertran et al., , 2021Haase et al., 2007;Lehmkuhl et al., 2021). In contrast, in Southern Europe (i.e. in the Iberian Peninsula, Southern France and along the Adriatic coast), loess deposits have a smaller extent and are mostly discontinuous (Costantini et al., 2018;Coudé-Gaussen, 1990). In these areas, smaller glacial systems (Pyrenees, Iberian System, Dinarides) are believed to be responsible for feeding rivers (Bertran et al., 2021;Wolf et al., 2019). The mass of available materials that form these loess deposits is largely due to glacial abrasion processes.
It was much more active in areas covered by large masses of ice, which later became the sources of particulate materials (Bertran et al., 2021).
Recent studies propose several genetic models based on the climatic and geomorphological characteristics of loess source areas (Lehmkuhl et al., 2021), especially the abrasion capacity of glacial systems and the type of connection with subsequent fluvial and aeolian transport processes (Bertran et al., 2021;Li et al., 2020). All these models consider combined transport systems that start in glacial environments as the main mechanisms to produce fine particles of sand and silt, which would later be taken up and reworked by river drainage networks. They would finally be transported by winds from alluvial areas towards annexed landscapes at a regional or continental scale. This interpretation has led to the idea that the alluvial plains of large rivers are amongst the most appropriate systems for acting as potential source areas of wind-transported particles Muhs, 2007;Smalley et al., 2009).
Although there are some precedents (Solé-Sabarís et al., 1957), loess deposits of the Iberian Peninsula were not described and analysed in depth until a few decades ago. The most important loess assemblies correspond to the large Cenozoic basins of the Ebro River (Boixadera et al., 2015;Iriondo & Kröhling, 2004;Poch et al., 2021) and the Tagus River (Calvo et al., 2016;Garcia-Giménez et al., 2012;Wolf et al., 2019). In the Ebro and Tagus loess areas, the average grain size ranges from coarse silt to fine sand. They comprise accumulations 2-8 m thick, with discontinuous and irregular outcrops exhibiting high carbonate contents (35-60%). Extension and location of the deposits are influenced by aerodynamic factors related to regional orography and katabatic wind adaptation. Other sets of loess deposits have been described at scattered locations on the Iberian Peninsula (La Mancha, Girona), but they are much smaller entities than the previous ones (Mücher et al., 1990).
The relationships between fluvial network, regional wind systems and loess accumulations will help us to know the dust provenance areas, main transport direction, travel distance and orographic effect of surrounding landscapes. Hence, the paleogeographic and paleoenvironmental reconstruction of the Ebro Valley would allow a better understanding of climatic and geomorphological dynamics (fluvial, alluvial, aeolian, weathering) operating in the region during the last phases of the Quaternary.
To establish the loess source areas, various techniques have been developed based on the recognition of specific or tracing characteristics in both sediments and source areas. The techniques consisted of the identification of heavy minerals (Chmielowska & Salata, 2020;Muhs et al., 2018;Peng et al., 2016;Tham o-Bozs o et al., 2014;Wolf et al., 2019), geochemical composition (Bosq et al., 2020;Schatz et al., 2015;Waroszewski et al., 2021) and U-Pb isotope analysis of zircons (Fenn et al., 2022;Újvári et al., 2016;Wolf et al., 2021). One of the most useful and effective methods is the comparison of sets of heavy minerals present in the loess sand fraction and their potential source materials: fluvial terraces and regional lithological substrates (Chmielowska & Salata, 2020;Peng et al., 2016;Tham o-Bozs o et al., 2014;Wolf et al., 2021). As ancillary techniques, particle grain size analysis of the loess and original materials, and analyses of the general chemical composition, are also used. The main criteria employed in the interpretation of the possible origin of aeolian deposits are: (a) the average size of the grains is related to the mode of transport-saltation, suspension at low or high altitude and the distance of transport, and understanding that coarse fractions correspond to nearby (local) contributions; and (b) the mineralogical nature of particles, in which changes in composition can be interpreted as changes in wind direction or in the source of deflated materials (Wolf et al., 2019). Recent studies on loess origin (Bosq et al., 2020;Waroszewski et al., 2021;Wolf et al., 2019) have found that the alluvial plains of large rivers and the sedimentary substrate and soils of interfluvial areas are the preferential zones for the mobilization of loess particles (Römer et al., 2016;Smalley et al., 2009).
Previous studies on loess of the Eastern Ebro Valley (Boixadera et al., 2015;Plata et al., 2021) show coarse sizes of loess particles indicating proximal provenance. Our research aimed to identify source areas of the aeolian materials from Zaragoza to the east, the prevailing wind direction during loess deposition, possible transport distances and the role of geographical features in the separation of depositional areas in the Ebro Valley.
The working hypothesis was based on accepting three assumptions: (a) source areas are relatively close when grain sizes are large (Pye, 1995); (b) the mineralogical signature of loess is inherited from that of the wind-transported sediments and thus can be determined with techniques of separation and identification of heavy minerals (Tham o-Bozs o et al., 2014); and (c) alluvial plains of large rivers may be a potential source of particles Smalley et al., 2009). Techniques of cartography and field description, particle size analysis, mineralogical separation and identification were combined to carry out our research.

| STUDY AREA
The Ebro River Valley, located in the NE of the Iberian Peninsula with a drained area of about 85 000 km 2 , is one of the largest basins in the Western Mediterranean. The triangular shape of the river basin corresponds almost exactly to the former Cenozoic foreland basin located between the Pyrenees, the Iberian Range and the Catalan Coastal  (Arche et al., 2010;García-Castellanos et al., 2003) to the Pliocene (Babault et al., 2006). Sediments of the centre of the Ebro Valley are basically Miocene evaporitic sequences in the western zone (evaporites around Zaragoza), whereas they are detrital series of Oligocene and Miocene sandstones, marls, lutites, lacustrine limestones and more rarely evaporites in the eastern zone between Sástago and Mequinenza (Solà & Costa, 1997) (Figures 1 and 2). The Middle Ebro (Ollero, 1996)  The bottom of the Ebro Valley ( Figure 2) has well-developed Quaternary river terrace levels with a maximum width of 10 km. Downstream of Sástago, the course begins to have a noticeable increase in meanders, 400-600 m wide, and it incises into the valley, forming canyons. The Cinca and Segre rivers have a 2-3 km-wide terrace system. The M ora Basin (Figures 1 and 2) is a small graben located in the Catalan Coastal Ranges, near the contact with the Ebro Basin delimited by normal faults active during the Neogene (Teixell, 1988).
The climate is Continental Mediterranean, with little rainfall and extreme temperatures that make the central area of the basin (Los Monegros) the northernmost arid area in Europe (Valero-Garcés et al., 2005). Annual rainfall can reach 1500 mm in the Pyrenees but drops sharply to <400 mm in the centre of the valley, due to the shadow effect of the Iberian Range. Annual evapotranspiration can exceed 1200 mm. The winds are dry and strong with a WNW-ESE direction, locally known as 'cierzo' (Gutiérrez et al., 2013;Mensua & Ibáñez, 1975;Puicercús et al., 1997).
The climate underwent major changes in the region during the Quaternary, with the alternation of cold and warm climates (González-Sampériz et al., 2008). On a global scale, precipitations (P) were lower during the glacial periods because of lower sea temperatures. However, evapotranspiration (E) was also weaker on the continent. Over most of Spain, climate modelling suggests higher soil moisture during the glacial period of the Last Glacial Maximum because of the much higher P-E budget (Scheff et al., 2017;Strandberg et al., 2011). These changing environments led to a succession of complex morphogenetic systems. In cooling periods, glacial and periglacial processes dominated in mountain areas and a combination of alluvial aeolian environments were developed at lower F I G U R E 1 Geological map of the Ebro Valley. The location map (a) shows the Ebro Valley in green and the study area in yellow. The main map (b) is derived from Pardo et al. (2004) and the geological map of Spain 1:1 000 000 from Rodríguez et al. (2015).  (Lehmkuhl et al., 2021) and considered as small continental systems in Cenozoic basins (Bertran et al., 2021). Their dominant grain size fraction is very fine sand followed by coarse silt, so they have been classified as sandy loess in the USDA textural classification (Bertran et al., 2021;Coudé-Gaussen, 1990). Their average mineral composition is dominated by quartz (40-45%) and calcite (28-45%).
Organic matter is very scarce (average 0.35%). Deposits are usually 2-5 m thick, although in very favourable positions they can reach 10 m.
Their high lateral discontinuity makes them very difficult to map at regional scales. Most of the deposits correspond to very young sequences, dated between 17 and 34 ka with OSL techniques, that have been related to cooling phases of the MIS-2 stage. Occasionally, older complex sequences have been identified in Mas de l'Alerany and Chiprana profiles, with ages between 130 and 144 ka with the post-IRSL method, and thus they would be related to stage MIS-6 (Boixadera et al., 2015;Dominik et al., 2021;Plata et al., 2021).

| METHODOLOGY
Identifying the provenance of the aeolian materials that form the loess deposits of the Eastern Ebro Valley has been based on the statistically tested similarity of (1) the grain size, (2) the mineralogical signature and (3) the chemical properties between the source area and the loess deposits. In the case of grain size, the degree of particle size sorting and fractioning caused by the wind deflation and transport plays a major role (Bertran et al., 2021;Bosq et al., 2018;Crouvi et al., 2008).
The heavy mineral analyses compare the presence and frequency of representative mineral species in sources and deposits (Chmielowska & Salata, 2020;Römer et al., 2016;Song et al.,  gypsum contents have also been selected as common chemical components. Their presence in loess deposits can be attributed to a potential contribution of Miocene interfluvial lithologies. Gypsum in loess can be related to the evaporitic sequences of Miocene formations near Zaragoza as a major source . We analysed the grain size distribution of 47 samples of the potential source areas: 42 from fluvial terraces and 5 from Miocene substrates (Supplementary Table S1). The fluvial sediments were collected along 350 linear km of Ebro, Cinca and Segre river terraces ( Figure 2). These terraces are 8-20 m higher than the present-day talweg, and have ages around 47 AE 4 ka, obtained by correlation from the dated Cinca and Segre lower terrace levels (Lewis et al., 2009(Lewis et al., , 2017Stange et al., 2013); therefore, they would correspond to the floodplain levels contemporary with the formation of the loess deposits (Boixadera et al., 2015;Plata et al., 2021). The Miocene substrate samples consist of sandstones, limestones and marls, representative of the main lithostratigraphic units located east of Zaragoza ( Figure 2). Potential Miocene sand sources correspond to sandstone paleochannels with a much reduced surface outcropping.
Regarding the loess deposits, 195 samples were collected (Supplementary Table S1). Among them, 175 correspond to systematic samplings of 10 vertical profiles with depth intervals from 20 to 50 cm. An additional 20 samples were taken at 16 single points to obtain a more complete spatial distribution. Figure 3 shows examples of the sampled terraces and loess sediments.
The grain size analyses were carried out with a Coulter 230 laser diffractometer, which has a resolution of diameters between 0.04 and 2000 μm in the University of Barcelona's Laboratory of Sedimentology. The samples were previously treated with H 2 O 2 to remove organic matter and with sodium polyphosphate, which acts as a dispersant, for 6 h. The percentages of sand, silt and clay (USDA classification) were plotted in the USDA textural triangle.
The identification of heavy mineral composition and the analyses of chemical properties (calcium carbonate and gypsum contents) were carried out on 18 alluvial terrace samples (10 from the Ebro River and 8 from the Cinca and Segre tributaries). Only two samples of the Miocene substrate contained sufficient heavy mineral content for their analysis. Besides, 38 samples were collected from loess deposits (22 from the different profiles and 16 from the single points). A summary of the samples is presented in Supplementary Table S1.
The heavy mineral analysis consisted of a pretreatment of 60 g of sample to destroy the organic matter and the particulate aggregates, as in the case of grain size analysis. To avoid coatings and aggregates, they were further subjected to an ultrasonic bath with a solution of nitric acid. The fraction size 50-250 μm was selected for the determination of heavy minerals, since it coincides with the most representative loess size in the region, and because this grain size facilitates their determination through the microscope (Morton & Hallsworth, 1994).
The identification of heavy minerals was conducted on the 50-250 μm fraction, after being attacked with 50% HCl to remove carbonates and washed repeatedly with deionized water to extract remaining reagents. About 5-10 g of the fraction underwent a double gravimetric separation of heavy minerals with dense liquids. Two decantation phases were performed: the first 10 min with bromoform (CHBr 3 , density 2.89 g/cm 3 ) and the following 20 min with diiodomethane or methylene iodide (CH 2 I 2 , density 3.32 g/cm 3 ). The separation process was undertaken using a procedure similar to that described by Mange & Mauer (1992) and Fernández (2016).
Light minerals float in bromoform, whilst heavy minerals (HM) sink. Very heavy minerals (VHM), heavier than 3.32 g/cm 3 , were also determined as those sinking into the diiodomethane. This gravity method has been used successfully in previous mineral studies (Fenn et al., 2022;Gonçalves et al., 2017;Lin et al., 1992;Morton, 1985;Takehara et al., 2018). VHM include zircon, augite, diopside, staurolite, rutile, barite, garnet group, epidote and kyanite, these being the most representative. To simplify the description, pyroxenes and amphiboles were grouped as inosilicates. The diiodomethane method normally selects about 60% of the mineral species separated with bromoform, and discriminates rock fragments and micas, thus clearly facilitating mineral identification and counting (Mange & Mauer, 1992).
The VHM fraction was mounted on thin sections and fixed with resin. The identification of VHM species was performed with a petrographic polarized light microscope following the manuals of Mange & Mauer (1992) and Andò et al. (2012). In parallel, identification of minerals with a binocular microscope was conducted following the descriptions of Fernández (2016) and the website Sandatlas (2021).
The number of identified specimens (grains) was counted following a total counting method as indicated by Mange & Mauer (1992) and Bars o (2006). The counting surface used was 1 cm Â 1 cm plots for the source areas and 0.5 cm Â 0.5 cm plots for the loess deposits, since the latter presented thinner grains and were consequently more concentrated, with a minimum of 150 transparent specimens per sample.
Results were calculated as abundance percentages of (a) heavy minerals (%HM), (b) very heavy minerals (%VHM), (c) transparent minerals in VHM fraction (%Trans) and (d) 17 VHM species identified in the mineral set.
Calcium carbonate was determined by Bernard calcimetry, with an attack of 50% HCl on 0.5 g of sample of the 50-250 μm fraction following the process described in Porta et al. (1986). Gypsum content was obtained by the gravimetric method from the dehydration of gypsum by heating samples in an oven at 105 C (Artieda, 2013;Artieda et al., 2006) and turbidimetric determination using BaCl 2 .
The statistical analyses were performed on 22 variables: percent- Pearson correlation tests with a p-value <0.05 were considered significant. Subsequently, a Kruskall-Wallis univariate analysis (H-test) was carried out on the seven most outstanding variables of the PCA, to know whether the different samples were equidistributed and therefore whether they belonged to the same population. If the differences were significant according to Kruskall-Wallis, we applied Dunn's test to perform pairwise comparisons between each independent group, in order to know which groups were significantly different. The statistical analyses were carried out using the statistical program PAST v.1.89 (Hammer et al., 2001).

| Exploratory analyses of the source and loess areas
The samples from source areas and loess deposits were subdivided into tentative groups considering their geomorphological setting. The potential source areas consist of fluvial terraces and interfluves on Miocene materials, subdivided as follows. (1)  Although the cumulative variance of the first two axes is not very high, some groupings are observed that are consistent with previous tentative groups (Figure 4). The first axis divides both the sections of the terraces and the loess areas. In the case of the terraces, it locates the ME to the right part of the graph, with higher percentages of zircon, staurolite and rutile, and the C + S and LE on the left part, with higher percentages of transparent minerals and inosilicates. The C loess samples have similar characteristics to the ME, but differ from the MB and EC samples, which can be grouped with C + S plus LE. The second axis separates the loess samples from the terraces (lower and upper side of the graph, respectively), the latter having higher mean/mode grain sizes and higher percentages of heavy minerals than those of loess (Figure 4) The Kruskall-Wallis test showed significant differences (p < 0.05) between the samples of sediment sources for all the variables analysed, specifically between the Middle Ebro and the Cinca-Segre system, and between the loess from the central zone and the loess of M ora Basin (Table 1).

| Grain size
The average particle size of the fine earth (<2000 μm) on the source areas is similar, although the mode of the grain size of Cinca and Segre systems is higher than ME and LE. The average diameter of loess grains was clearly smaller than that from the terraces. Loess from C (right bank of the Ebro River) and MB are the coarsest ones, while the other marginal areas (NC, SC, EC) decrease slightly, but only significantly in the NC. The mode results show similar behaviour to the mean ( Table 2).
The texture triangle ( Figure 5) shows that the sand fraction of the terraces ranges from 60% to 90% and in loess from 20% to 70%. The sand contents of C and MB deposits and C + S terraces have intermediate values between 50% and 70%. The coarse texture of the loess allows us to classify them as sandy loess (37-67% sand; Bertran et al., 2021) and as wind-borne sands in the few cases where sand content exceeds 67%. The Ebro Basin loess decrease in particle size from the central zone to the surrounding areas. The grain size of the terraces is larger than that of the loess; therefore, the hypothesis of the terraces being the source of the loess should be considered. Figure 6 and Table 1 show the distributions of heavy minerals in the 50-250 μm fraction separated by bromoform, sampled from source areas and loess deposits. There are significant differences depending on the river reach considered (ME, C + S and LE): the percentage in the ME was 2.36 AE 0.46%, while in the Cinca and Segre tributaries it was fourfold (C + S, 10.96 AE 2.76%). As a result of this enrichment, the LE increased its content to 5.20 AE 2.77%, although differences between ME and LE were not significant according to Tukey's Q test.

| Total heavy mineral percentage (%HM)
In general, Miocene sediments from the interfluves (MI) contained a much lower percentage of heavy minerals than the fluvial systems, with an average of 0.19 AE 0.46% (except in sandstone formations, where they were 0.88 AE 0.04%). Therefore, the Miocene sediments acting as source areas would tend to reduce the percentage of heavy minerals. T A B L E 1 Percentage and standard deviation of heavy mineral analyses. Values followed by the same letter/colour are not significantly different (p < 0.05) within each group (source areas or loess deposits). In the case of Miocene, the five samples are used to obtain the percentage of heavy minerals (%HM and %VHM) but only two samples have enough minerals to perform the analysis.

Miocene samples are not included in the Kruskal-Wallis test. S is the number of samples. Original data in Supplementary
In the loess deposits ( Figure 6 and

| Very heavy mineral composition (VHM)
The most representative very heavy minerals studied in the source areas and loess deposits were zircon, staurolite, rutile and the inosilicates group (Table 1, Supplementary Table S1). The other minerals analysed (garnet, epidote, kyanite, monazite, barite, spinel, corundum, anatase, zoisite, titanite) were unrepresentative or did not show a differentiated spatial pattern to make them useful as tracer minerals. Miocene formations would be considered a secondary source of loess sediments. Generally, they had a rather low content of VHM.

| Source areas
They contained some minerals from the Iberian Range denudation, such as zircons or garnets, but the low number of samples (2) did not allow statistical analyses. Figure 9 and Table 1 show the percentages of different mineral species (%VHM) identified in loess. In C, NC and SC, the sum of zircon, staurolite and rutile tends to be higher than the inosilicates, while in MB and EC the inosilicates are more important. The rest of the minerals have a low representation or do not show sufficient contrast between areas, as happened in the terraces; therefore, they are not considered as tracers of source areas.

| Loess
Zircon is the mineral with the highest contrast in the spatial distribution ( Figure 8). In the zone of the greatest accumulation (C), zircon accounts for 29.84 AE 4.09%, significantly decreasing away from the F I G U R E 7 Histogram of the mineral species obtained with diiodomethane separation in the source areas, formed by the terraces of the Middle Ebro River (ME), Cinca-Segre (C + S), Lower Ebro River (LE) and Interfluvial Miocene formations (MI). Original data can be found in Supplementary Table S2

| Gypsum and carbonates
Gypsum does not appear in the terrace sands due to its high solubility.
Nevertheless, it is ubiquitous in the Miocene sediment outcropping in the interfluves of the Ebro Basin . In some lithologi-

| DISCUSSION
The Eastern Ebro Valley loess have a very coarse mean (40-52 μm) and mode (50-79 μm) grain size, comparable with that of sandy loess in Belgium (Bertran et al., 2021) and Tunisia (Coudé-Gaussen, 1990;Faust et al., 2020). These coarse textures suggest very proximal source areas (Pye, 1995), mainly related to fluvial alluvial plains (Fenn et al., 2022;Lehmkuhl et al., 2016;Smalley et al., 2009 Therefore, the provenance of this loess is mainly the Lower Ebro (LE). River. They are rich in carbonates and gypsum, but poor in VHM (zircons, staurolite, inosilicates). Fine particle availability to wind deflation is clearly higher in terraces than in Miocene substrate interfluves (sandstone, limestones, marls), where very low rock weathering rates and vegetation cover result in low sediment supply and availability (Bullard et al., 2008(Bullard et al., , 2011. In the Ebro depression, a 50% reduction in %HM took place between the source area (ME) and the depocenter (C The similar mineralogical composition of the oldest and more recent sequences in Mas de l'Alerany and Chiprana, attributed to a MIS-6 phase and a MIS-2 phase, respectively, suggests that there were no substantial changes in the main source of sediments during this time interval and that the direction of prevailing efficient winds did not differ either (Gutiérrez et al., 2013;Mensua & Ibáñez, 1975;Plata et al., 2021;Schatz et al., 2015;Wolf et al., 2019).
Terraces of the Middle Ebro have a particle deflation level Regarding the methodology used in mineralogical analysis, the use of diiodomethane is revealed as a useful technique to separate and identify VHM signatures of loess and source areas, because it shows a good correlation with the classic bromoform separation techniques and reduces noise from rock fragments and micas.
Besides, it would be interesting to increase the number of samples to obtain more significant results, especially in the EC and SC distal areas.

| CONCLUSIONS
The significant loess deposit cover in the Eastern Ebro Valley is formed by fine sands and silts with a grain size mode between 50 and 79 μm. They are coarser than the classic loess and this implies a closer source area.
Textures and particle mineralogical signatures of the Ebro, Cinca and Segre rivers, as well as of interfluve areas, are sufficiently contrasted to separate source areas and relate them to different groups of loess deposits. In particular, the proportion of HM and some mineral concentrations in loess (gypsum, zircon, staurolite, inosilicates) can be used as effective tracers for particle provenance.
Our results show two clearly differentiated deposition areas related to two major source areas. The first is the loess deposits of This publication is part of the I + D + i RTI2018-094927-B-I00 project, funded by the Spanish MCIN/AEI and FEDER 'A way to make Europe', and has received a grant for its linguistic revision from the Language Institute of the University of Lleida (2022 call).

DATA AVAILABILITY STATEMENT
Data are available in the online Supplementary Material.